Dolly Varden, overwintering, and the subsistence calendar.
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Dolly Varden feed at sea and return to overwinter in specific river reaches, which makes those reaches the whole population's bottleneck.
Subsistence harvest is organised around when the char are where, and that timing is the practical calendar of the region.
Chum salmon use the same systems and their northern distribution has been shifting, which is documented rather than anecdotal.
Elsewhere on the river